Holt has been with NBC News since 2000. He steps in for Brian Williams on the NBC Nightly News Monday night. For Holt, the circumstances are new and treacherous, but the job itself is not. He's been filling in for Williams for almost a decade and he's a friendly face in the halls of NBC News.
The 55-year-old newsman has been a co-host of the weekend editions of "Today" since 2003. He added weekend "Nightly News" to his duties in 2007, so he works pretty much nonstop Saturday and Sunday.
He also hosts the newsmagazine "Dateline."
The nonstop nature of Holt's work once garnered him the nickname "Iron Pants."
He joined NBC in 2000 after two decades in local TV. He was one of the main anchors on NBC's cable news channel, MSNBC, during the start of the Iraq War in 2003.
"Lester is always pitch perfect in the anchor chair," said Mark Effron, a former vice president of daytime at MSNBC. "You can't throw him a curve he can't take."
Effron, who is now a media consultant and adjunct journalism professor at William Paterson University, said that Holt never begrudgingly took an assignment while at MSNBC.
This week is, in some ways, his most challenging assignment yet. With the crisis swirling around Williams, Holt has the difficult task of steadying the ship and maintaining Williams' top-rated audience.
At the same time, he can't get too comfortable in the job, lest he appear to be taking advantage of the situation.
Back in 2013, when Williams was undergoing knee surgery, Holt filled in on weekdays and held NBC's top spot, bringing in 7.6 million viewers the week before Williams' return.
Some observers have cited those ratings to say that Holt will be well-received this week.
Indeed, Holt was an obvious choice when Williams decided on Saturday to, in his words, "take myself off" the weekday "Nightly News" for "the next several days."
"Lester Holt has kindly agreed to sit in for me," Williams said.
Related: Will Brian Williams return?
Holt acknowledged the awkwardness on Monday night, saying "we want to take just a moment to tell you where Brian is tonight," and then describing Williams' announcement from Saturday.
"He'll be off while this issue is dealt with," Holt added.
The announcement of Holt as Williams' sub hushed speculation -- at least momentarily -- about who might step in for Williams either temporarily or permanently. Other names have included "Today" co-hosts Matt Lauer and Savannah Guthrie and a new arrival from ABC, Josh Elliott.
Certainly someone like Lauer is higher-profile than Holt, and that has led some TV critics to wonder if Holt is right for the weekday post over the long term.
"The position of broadcast evening news anchor is still important enough that it calls for an extra level of telegenic presence, some intangible dash of starpower," the New York Daily News TV critic David Hinckley wrote Monday. "That's probably not Holt."
Others have argued that someone like Holt is perfect precisely because he doesn't have as much starpower as Williams.
Related: Brian Williams controversy: Here's what we know
It's unclear how long Holt will be filling in, since Williams' leave of absence has no end date.
Some inside NBC are openly wondering if Williams will be able to return. And in the meantime, Holt is there.
"However long he's in that seat, he will represent the very best principles of journalism," Effron said. "There's nobody better positioned if that job were to open up than Lester Holt."
CNNMoney (New York) February 9, 2015: 7:09 PM ET
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